James and Sadie Goodenough have settled where their wagon got stuck – in the muddy, stagnant swamps of 1830s Ohio. They and their children work relentlessly to tame their patch of land, buying saplings from a local tree man known as John Appleseed so they can cultivate the fifty apple trees required to stake their claim on the property. But the orchard they plant sows the seeds of a long battle. James loves the apples, reminders of an easier life back in Connecticut; while Sadie prefers the applejack they make, an alcoholic refuge from brutal frontier life.
Fifteen years later, the youngest Goodenough, Robert, is wandering through Gold Rush California. Restless and haunted by the broken family he left behind, he has made his way alone across the country. In the redwood and giant sequoia groves he finds some solace, collecting seeds for a naturalist who sells plants from the New World to the gardeners of England. But you can run only so far, even in America, and when Robert’s past makes an unexpected appearance he must decide whether to strike out again or stake his own claim to a home at last.